Brief CV

Anne Chao (趙蓮菊)

Address: Institute of Statistics

National Tsing Hua University

Hsin-Chu, Taiwan 30043

Tel/Fax: 886-3-5715131 ext 33161

886-3-5728318 (fax)


Google Scholar

Research Gate

Education 1973                 B. S. (Mathematics), National Tsing Hua University, Taiwan

1977                 Ph.D. (Statistics), University of Wisconsin-Madison, USA

Experiences 1977-78            Visiting Assistant Professor, Department of Statistics University of  Michigan-Ann Arbor, USA

1978-82            Associate Professor, National Tsing Hua University

1985-86            Visiting Scientist, Department of Biostatistics, Harvard University, USA

1982-present     Professor, National Tsing Hua University

Major Professional Services: 1994-2013       Associate Editor, Biometrics

1998-2011       Associate Editor, Journal of Agricultural, Biological, and Environmental Statistics

1991-96, 1999-2005 Associate Editor, Statistica Sinica

2001-2009       Associate Editor, Australian & New Zealand Journal of Statistics

2015-              Associate Editor, Methods in Ecology and Evolution

Referees for various journals in biology, ecology, statistics, epidemiology and related sciences.

Honors: 1990-95        Taiwan National Science Council Award for Outstanding Research (傑出研究獎三次)

1987             Elected Member of International Statistical Institute

1996             Best Teaching Award, National Tsing Hua University (傑出教學獎)

1997             Fellow, Institute of Mathematical Statistics

2001             ISI Citation Classic Award for two highly-cited papers (Chao, 1984; Chao and Lee, 1992)

2003             Academic Award, Ministry of Education, Taiwan (教育部學術獎)

2005-2008     Taiwan National Chair Professor (國家講座)

2006-            Tsing Hua Distinguished Chair Professor (清華特聘講座)

Theses Supervised: ~ 100 Master Theses;  22  Ph.D. Theses

Ph.D Theses:

Lee, S-M (1990) Estimating population size for closed capture-recapture models via sample coverage. (Feng-Chia University)

Ma, M-C (1992) Stopping rule and estimation for recapture debugging with unequal detection rates. (National Cheng-Kung University)

Hwang, W. D. (1992) Quantifying the effects of unequal catchabilities on Jolly-Seber estimators. (National Cheng-Kung University)

Chu, W-T (1995) A unified martingale estimation approach to estimating parameters of time-specific model. (Feng-Chia University)

Chen, Y-C (1996) The estimation of population size for capture-recapture experiments under Pollock’s robust design. (Chia-Nan College of Pharmacy and Science)

Hwang, Wen-Han (1997) Estimating population Size for continuous time capture-recapture experiments. (National Chung-Hsin University)

Tsay, P-K (1997) Comparisons of loglinear model and sample coverage methods in capture-recapture experiments. (Chang-Kang University)

Lin, S-P (2001) Comparison of population size estimation between continuous and discrete capture-recapture experiments. (Chung-Hua University)

Yang, H-C (2002) The applications of Markov chain models and kernel smoothing in capture-recapture experiments. (Institute of Statistical Science, Academia Sinica)

Shen, T.- J. (2003) Prediction of Biodiversity. (National Chung-Hsin University)

Li, P.-C. (2005) Species estimation and similarity indices in quadrat sampling. (Wistron NeWeb Co. Industrial Park, Hsin-Chu)

(Co-advisee) Lee, P.-L. (2006) Clustering for longitudinal data. (Department of Statistics, Tamkang University)

Chiang, S.-C. (2006) Similarity indices for multiple communities. (Biostatistics Task Force, Taipei Veterans General Hospital)

Pan, H. Y. (2007) Estimation of shared species richess in multiple communities. (National Chia-Yi University)

Han, R.-P. (2008) Statistical estimation of beta diversity. (Central Taiwan University of Science and Technology)

Jiang, Y.-H. (2008) Multiple-community similarity indices under both discrete and continuous diversity data.

Lin, C.-W. (2009) Estimation of biodiversity in sampling without replacement. (Chaoyang University of Technology)

Lin, S.-Y. (2012) Hierarchical and space-time decomposition of biodiversity indices.

Wang, Y.-T. (2013) Statistical estimation of Hill numbers and similarity indices.

Hsu, F.-J. (2013) Statistical estimation of rarefaction and extrapolation curve for beta diversity.

Hsieh, T.-C. (2014) Rarefaction and extrapolation with biodiversity.

Ma, K.-H. (2016) Rarefaction and extrapolation of similarity/differentiation measures and species diversity in sampling without replacement.

Major Research Area: Ecological statistics and bio-diversity estimation

  • Species estimation and population size estimation
  • Animal abundance models and estimation
  • Capture-recapture models and their applications
  • Bio-diversity indices (species, phylogenetic and functional diversities) and their estimation
  • Biological sampling methods
Publications: 所出版論文被SCI期刊作者引用 (Web of Science) 次數已超過 9750 次, Google Scholar超過 18200次(Accessed on May 8, 2018)

A full list of publications is given on the website.


  • Lou Jost and Anne Chao: “Diversity Analysis”, Chapman & Hall, London. (Under Contract, Expected publication time 201?)
  • Anne Chao, Chun-Huo Chiu and Lou Jost: “Statistical Estimation of Biodiversity Indices”, John Wiley & Sons. (Under Contract, Expected publication time 201?)
Software and User’s Guide:

(freely downloadable from

  • CARE (CApture-REcapture) including

          CARE-1: for capture-recapture epidemiological data

          CARE-2: for discrete-time capture-recapture models

          CARE-3 : for continuous-time capture-recapture models

          CARE-4 : for Markov chain capture-recapture models

  • SPADE (Species Prediction And Diversity Estimation): for biodiversity data analyses
  • iNEXT (Interpolation/EXTrapolation)
  • iNEXT-pd (Interpolation/EXTrapolation for phylogenetic diversity)
  • PhD (Phylogenetic Diversity)